1977-1983 Medical and graduate student, Temple University 1983 Postdoctoral Fellow with Dr. Paul Ts'o, Division of Biophysics, School of Hygiene and Public Health, The Johns Hopkins University 1983-1989 Postdoctoral fellow with Dr. David Agard, Department of Biochemistry and Biophysics, University of California, San Francisco 1989-1995 Assistant Professor, Department of Cell and Structural Biology, University of Illinois, Urbana 1995-1999 Associate Professor, Department of Cell and Structural Biology, University of Illinois, Urbana 1999-2005 Professor, Department of Cell and Structural Biology University of Illinois, Urbana 1/04 - 6/04 Interim Head, Department of Cell and Structural Biology 2005-present Professor, Department of Cell and Developmental Biology University of Illinois, Urbana 1/2008-1/2014 Head, Department of Cell and Developmental Biology University of Illinois, Urbana